In 2200, private detective Aline Ruby and her android partner Carlos Rivera are hired by a wealthy businessman to track down a notorious hacker. On Mars, they descend deep into the underbelly of the planet's capital city where they uncover a darker story of brain farms, corruption, and a missing girl who holds a secret about the robots that threatens to change the face of the universe.
Miraculous holders from another world appear in Paris. They come from a parallel universe where everything is reversed: the holders of Ladybug and Black Cat Miraculouses, Shadybug and Claw Noir, are the bad guys, and the holder of the Butterfly Miraculous, Hesperia, is a superhero. Ladybug and Cat Noir will have to help Hesperia counter the attacks of their evil doubles and prevent them from seizing the Butterfly's Miraculous. Can our heroes also help Hesperia make Shadybug and Claw Noir better people?