As Pretty Cures all over the world fight against Saiarks, minions of the evil Phantom Empire led by Queen Mirage, Himelda "Hime" Window, also known as Cure Princess, feels downhearted after losing yet again to the Saiark, having to be bailed out by another Cure named Cure Fortune. Blue, the god of the universe, gives Hime a Crystal of Love so that she can find a partner to fight alongside her. Excited to be able to have a friend, Hime decides to randomly throw the crystal to decide her partner, which randomly lands on a girl named Megumi Aino. Hime, along with her fairy partner Ribbon, follows Megumi as she goes about her shopping, helping others when they're in trouble.
Megumi Nakajima
Megumi Aino / Cure Lovely (voice)
Megumi Han
Hime Shirayuki / Cure Princess (voice)
Rina Kitagawa
Yuuko Omori / Cure Honey (voice)
Haruka Tomatsu
Iona Hikawa / Cure Forture (voice)
Naoko Matsui
Ribbon (voice)
Miyuki Kobori
Glasan (voice)
Shouma Yamamoto
Blue (voice)
Ryosuke Kanemoto
Seiji Sagara (voice)
Mariko Kouda
Queen Mirage / Mirage / Cure Mirage (voice)
Kazuhiko Inoue
Deep Mirror / Red (voice)
Takehito Koyasu
Oresky (voice)
Akemi Okamura
Hosshiwa (voice)
Tetsuo Kanao
Namakelder (voice)
Hirofumi Nojima
Phanphan / Phantom (voice)
Takuya Masumoto
Saiarks, Choiarks